Describe the behavior of chromosomes during meiosisDescribe to move events during meiosisExplain the differences between meiosis and mitosisExplain the mechanisms within meiosis the generate genetic variation amongst the products of meiosis

Sexual reproduction requires fertilization, a union of 2 cells from two individual organisms. If those two cells every contain one collection of chromosomes, then the result cell has two to adjust of chromosomes. The number of sets the chromosomes in a cell is referred to as its ploidy level. Haploid cell contain one set of chromosomes. Cell containing 2 sets the chromosomes are called diploid. If the reproductive bicycle is come continue, the diploid cell must somehow minimize its variety of chromosome sets prior to fertilization can happen again, or there will be a constant doubling in the variety of chromosome to adjust in every generation. So, in addition to fertilization, sexual reproduction contains a nuclear division, known as meiosis, the reduces the number of chromosome sets.

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Most animals and plants are diploid, containing 2 sets of chromosomes; in each somatic cabinet (the nonreproductive cells of a multicellular organism), the nucleus contains two copies of every chromosome that are described as homologous chromosomes. Somatic cells are occasionally referred to together “body” cells. Homologous chromosomes space matched pairs containing gene for the exact same traits in identical locations follow me their length. Diploid organisms inherit one copy of every homologous chromosome from every parent; every together, castle are considered a full set of chromosomes. In animals, haploid cell containing a solitary copy of every homologous chromosome are uncovered only in ~ gametes. Gametes fuse with one more haploid gamete to create a diploid cell.

The nuclear department that forms haploid cells, which is called meiosis, is related to mitosis. Together you have learned, mitosis is part of a cabinet reproduction bicycle that results in identical daughter nuclei that are likewise genetically identical to the original parent nucleus. In mitosis, both the parent and also the daughter nuclei save on computer the same number of chromosome sets—diploid for many plants and animals. Meiosis employs numerous of the very same mechanisms as mitosis. However, the beginning nucleus is constantly diploid and the nuclei that result at the end of a meiotic cell division are haploid. To attain the reduction in chromosome number, meiosis is composed of one ring of chromosome duplication and two ring of atom division. Due to the fact that the events that occur during each the the division stages room analogous to the events of mitosis, the same stage names room assigned. However, because there space two rounds of division, the stages are designated with a “I” or “II.” Thus, meiosis i is the first round the meiotic department and is composed of prophase I, prometaphase I, and also so on. Meiosis ns reduces the variety of chromosome sets from two to one. The hereditary information is also mixed throughout this department to produce unique recombinant chromosomes. Meiosis II, in i beg your pardon the 2nd round of meiotic division takes location in a means that is similar to mitosis, has prophase II, prometaphase II, and also so on.

Interphase

Meiosis is came before by an interphase consists of the G1, S, and G2 phases, which are nearly identical come the phases preceding mitosis. The G1 step is the very first phase that interphase and also is concentrated on cabinet growth. In the S phase, the DNA the the chromosomes is replicated. Finally, in the G2 phase, the cell undergoes the last preparations for meiosis.

During DNA duplication of the S phase, each chromosome becomes created of 2 identical copies (called sisters chromatids) that are organized together at the centromere till they are pulled apart during meiosis II. In an pet cell, the centrosomes the organize the microtubules the the meiotic spindle additionally replicate. This prepares the cell because that the first meiotic phase.

Meiosis I

Early in prophase I, the chromosomes can be seen clearly microscopically. Together the nuclear envelope starts to rest down, the proteins connected with homologous chromosomes lug the pair near to every other. The chop pairing the the homologous chromosomes is called synapsis. In synapsis, the gene on the chromatids the the homologous chromosomes are specifically aligned with each other. One exchange the chromosome segments in between non-sister homologous chromatids occurs and is dubbed crossing over. This procedure is revealed visually ~ the exchange together chiasmata (singular = chiasma) (Figure 7.3).

As prophase ns progresses, the close association between homologous chromosomes begins to break down, and also the chromosomes proceed to condense, return the homologous chromosomes continue to be attached come each other at chiasmata. The number of chiasmata varies with the types and the length of the chromosome. In ~ the end of prophase I, the bag are organized together just at chiasmata (Figure 7.3) and also are called tetrads since the 4 sister chromatids of every pair that homologous chromosomes are currently visible.

The crossover occasions are the first source of genetic variation developed by meiosis. A solitary crossover event in between homologous non-sister chromatids leads to a mutual exchange of tantamount DNA between a maternal chromosome and a head chromosome. Now, when that sister chromatid is moved into a gamete, that will bring some DNA from one parental of the individual and also some DNA native the other parent. The recombinant sisters chromatid has actually a combination of maternal and paternal gene that did no exist before the crossover.

 
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Figure 7.3 In this illustration of the effects of cross over, the blue chromosome come from the individual’s father and also the red chromosome came from the individual’s mother. Crossover occurs between non-sister chromatids of homologous chromosomes. The an outcome is one exchange of hereditary material in between homologous chromosomes. The chromosomes that have actually a mixture of maternal and also paternal succession are referred to as recombinant and also the chromosomes the are completely paternal or maternal are referred to as non-recombinant.

 The an essential event in prometaphase i is the attachment of the spindle fiber microtubules to the kinetochore proteins at the centromeres. The microtubules assembled native centrosomes at opposite poles that the cell prosper toward the middle of the cell. In ~ the end of prometaphase I, each tetrad is attached to microtubules from both poles, through one homologous chromosome attached in ~ one pole and the various other homologous chromosome attached to the other pole. The homologous chromosomes space still organized together in ~ chiasmata. In addition, the atom membrane has damaged down entirely.

During metaphase I, the homologous chromosomes are arranged in the facility of the cell with the kinetochores dealing with opposite poles. The orientation of each pair of homologous chromosomes at the facility of the cabinet is random.

This randomness, dubbed independent assortment, is the physical basis for the generation the the second form of hereditary variation in offspring. Think about that the homologous chromosomes of a sexually reproducing biology are initially inherited together two separate sets, one from each parent. Using humans as an example, one set of 23 chromosomes is present in the egg donated through the mother. The father gives the other set of 23 chromosomes in the sperm the fertilizes the egg. In metaphase I, this pairs heat up at the midway suggest between the 2 poles that the cell. Since there is an equal chance that a microtubule fiber will encounter a maternally or paternally inherited chromosome, the setup of the tetrads at the metaphase key is random. Any maternally inherited chromosome may challenge either pole. Any paternally inherited chromosome may additionally face either pole. The orientation of every tetrad is live independence of the orientation of the other 22 tetrads.

In every cell that undergoes meiosis, the arrangement of the tetrads is different. The variety of variations relies on the variety of chromosomes consisting of a set. There are two possibilities because that orientation (for every tetrad); thus, the possible number of alignments amounts to 2n wherein n is the number of chromosomes every set. Humans have 23 chromosome pairs, which outcomes in over eight million (223) possibilities. This number does not encompass the variability previously created in the sisters chromatids by crossover. Offered these 2 mechanisms, the is extremely unlikely that any kind of two haploid cell resulting indigenous meiosis will have the same genetic composition (Figure 7.4).

To summarize the genetic aftermath of meiosis I: the maternal and also paternal genes space recombined through crossover events occurring on each homologous pair throughout prophase I; in addition, the random assortment of tetrads in ~ metaphase to produce a unique mix of maternal and also paternal chromosomes that will certainly make their method into the gametes.

 
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Figure 7.4 To show random, live independence assortment at metaphase I, take into consideration a cell v n = 2. In this case, there room two possible arrangements in ~ the equatorial plane in metaphase I, as presented in the top cell of every panel. This two possible orientations cause the manufacturing of genetically different gametes. With more chromosomes, the variety of possible arrangements rises dramatically.

 In anaphase I, the spindle fibers traction the connected chromosomes apart. The sister chromatids stay tightly bound with each other at the centromere. It is the chiasma connections that are damaged in anaphase I as the fibers attached to the fused kinetochores traction the homologous chromosomes apart.

In telophase I, the be separate chromosomes come at opposite poles. The remainder the the typical telophase events may or might not occur depending on the species. In part organisms, the chromosomes decondense and also nuclear envelopes kind around the chromatids in telophase I.

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Cytokinesis, the physical separation that the cytoplasmic materials into two daughter cells, wake up without reformation of the nuclei in other organisms. In practically all species, cytokinesis separates the cell materials by either a cleavage furrow (in animals and some fungi), or a cabinet plate the will eventually lead to formation of cell wall surfaces that separate the 2 daughter cells (in plants). At every pole, there is just one member of each pair that the homologous chromosomes, so just one full collection of the chromosomes is present. This is why the cells are taken into consideration haploid—there is just one chromosome set, even though there space duplicate duplicates of the collection because every homolog still is composed of 2 sister chromatids that room still attached to every other. However, back the sisters chromatids were once duplicates the the same chromosome, they room no longer similar at this stage because of crossovers.